1. Key Laboratory of Bio‐Resource and Eco‐Environment of Ministry of Education College of Life Sciences State Key Laboratory of Hydraulics and Mountain River Engineering Sichuan University Chengdu China
2. Institute of Molecular Plant Sciences The University of Edinburgh Edinburgh UK
3. College of Science Tibet University Lhasa China
4. Tibet Key Laboratory of Forest Ecology in Plateau Area of Ministry of Education Research Institute of Tibet Plateau Ecology National Key Station of Field Scientific Observation & Experiment of Alpine Forest Ecology System in Nyingchi Tibet Agriculture & Animal Husbandry University Nyingchi China