1. Pleistocene Helophorus (Coleoptera, Hydrophilidae) from Borislav and Starunia in the Western Ukraine, with a reinterpretation of M. Lomnicki's species, description of a new Siberian species, and comparison with British Weichselian faunas;Angus;Philosophical Transactions of the Royal Society of London Series B, Biological Sciences,1973
2. Evolutionary stability since the Pleistocene illustrated by reproductive compatibility between Swedish and Spanish Helophorus lapponicus Thomson (Coleoptera, Hydrophilidae);Angus;Biological Journal of the Linnean Society,1983
3. Spanish ‘endemic’Ochthebius as a British Pleistocene fossil;Angus;Latissimus,1993
4. Challenges and rewards in the identification of Pleistocene fossil beetles, with the description of a new species of Hydraena Kugelann (Coleoptera: Hydraenidae) from the Hoxnian Interglacial;Angus;Quaternary Proceedings,1997
5. The response of arctic Carabidae (Coleoptera) to climate change based on the fossil record of the Quaternary period;Ashworth;Annales Zoologici Fennici,1996