1. Key Laboratory of Coarse Cereal Processing, Ministry of Agriculture and Rural Affairs, Sichuan Engineering & Technology Research Center of Coarse Cereal Industrialization, School of Food and Biological Engineering Chengdu University Chengdu 610106 Sichuan Province China
2. Institute of Agricultural Research Tibet Academy of Agricultural and Animal Husbandry Sciences Lasha 850000 Tibet China
3. School of Preclinical Medicine Chengdu University Chengdu 610106 Sichuan Province China
4. Center for Agricultural Genetic Resources Research Shanxi Agricultural University Taiyuan 030031 China
5. Biotechnology and Nuclear Technology Research Institute Sichuan Academy of Agricultural Sciences Chengdu 610066 Sichuan Province China