1. Optics of Photosynthesis Laboratory Institute for Atmospheric and Earth System Research (INAR)/Department of Forest Sciences Viikki Plant Science Centre (ViPS) Faculty of Agriculture and Forestry University of Helsinki Helsinki 00014 Finland
2. Organismal and Evolutionary Biology (OEB) Viikki Plant Science Centre (ViPS) Faculty of Biological and Environmental Sciences University of Helsinki Helsinki 00014 Finland
3. Department of Forest Sciences Faculty of Agriculture and Forestry University of Helsinki Helsinki 00014 Finland