1. Institute of Science and the Environment, University of Cumbria, Ambleside Cumbria LA22 9BB UK
2. School of Biodiversity, One Health and Veterinary Medicine, College of Medical, Veterinary and Life Sciences University of Glasgow Glasgow G12 8QQ UK
3. School of Animal, Plant and Environmental Sciences University of the Witwatersrand Private Bag 3, Wits 2050 Johannesburg South Africa