1. CAS Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany Chinese Academy of Sciences Kunming 650201 China
2. Arkansas Biosciences Institute Arkansas State University Jonesboro 72401 Arkansas USA
3. School of Integrative Plant Science, Section of Plant Biology and the L.H. Bailey Hortorium Cornell University Ithaca 14853 New York USA
4. BTI Computational Biology Center Boyce Thompson Institute Ithaca 14853 New York USA
5. CAS Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture, Chinese Academy of Sciences Wuhan Botanical Garden Wuhan 430074 China
6. Department of Organismic and Evolutionary Biology, Herbaria Harvard University Cambridge 02138 Massachusetts USA