1. Key Laboratory of Wetland Ecology and Environment Northeast Institute of Geography and Agroecology Chinese Academy of Sciences Changchun, Jilin China
2. Biology Department San Diego State University San Diego CA USA
3. College of Earth Sciences Jilin University Changchun, Jilin China
4. College of Surveying and Mapping Engineering Heilongjiang Institute of Technology Harbin China
5. Key Laboratory of Forest Ecology and Management Institute of Applied Ecology Chinese Academy of Sciences Shenyang China
6. State Key Laboratory of Vegetation and Environmental Change Institute of Botany Chinese Academy of Sciences Beijing China