1. State Key Laboratory of Agricultural Microbiology Huazhong Agricultural University Wuhan China
2. The Provincial Key Lab of Plant Pathology of Hubei Province, College of Plant Science and Technology Huazhong Agricultural University Wuhan China
3. Key Laboratory of Environment Change and Resources Use in Beibu Gulf, Ministry of Education Nanning Normal University Nanning 530001 China
4. United States Department of Agriculture, Agricultural Research Service Washington State University Pullman WA 99164, USA