1. State Key Laboratory of Urban and Regional Ecology, Research Center for Eco-Environmental Sciences; Chinese Academy of Sciences; Beijing 100085 China
2. University of Chinese Academy of Sciences; Beijing 100049 China
3. Key Laboratory of Forest Ecology and Environment, State Forestry Administration; Institute of Forest Ecology, Environment and Protection, Chinese Academy of Forestry; Beijing 100091 China
4. Faculty of Veterinary and Agricultural Sciences; The University of Melbourne; Parkville Victoria Australia