1. Institute of Infection and Global Health, Faculty of Health and Life Sciences; University of Liverpool; Liverpool UK
2. Department of Microbiology; National Veterinary Institute; Uppsala Sweden
3. CIRAD, UMR ASTRE; Montpellier 34398 France
4. CIRAD, UMR ASTRE; Sainte-Clotilde La Réunion 97490 France
5. Health Protection Research Unit in Emerging and Zoonotic Infections; Liverpool L69 3GL UK
6. Institute of Integrative Biology, Faculty of Health and Life Sciences; University of Liverpool; Liverpool UK