1. Geomicrobiology Group; School of Life Sciences; University of Dundee; Dundee DD1 5EH Scotland UK
2. Electron Microscopy; Central Imaging Facility; Centre for Advanced Scientific Technologies; School of Life Sciences; University of Dundee; Dundee DD1 5EH Scotland UK
3. Laboratorio Biodiversità dei Funghi; Dipartimento di Biologia Ambientale; Sapienza Università di Roma; Rome 00185 Italy
4. The James Hutton Institute; Aberdeen AB15 8QH UK
5. Department of Soil and Environment; Swedish University of Agricultural Sciences; Uppsala Sweden
6. Laboratory of Environmental Pollution and Bioremediation; Xinjiang Institute of Ecology and Geography; Chinese Academy of Sciences; Urumqi 830011 China