1. UPMC University Paris 06; UMR 7621, LOMIC, UMS 2348, Observatoire Océanologique, Banyuls/mer F-66650 France
2. CNRS; UMR 7621, LOMIC, Observatoire Océanologique Banyuls/mer F-66650 France
3. Department of Marine Sciences; University of Georgia; Athens GA 30602 USA