1. Department of Marine Microbiology and Biogeochemistry, NIOZ Royal Netherlands Institute for Sea Research; and Utrecht University; Den Burg P.O. Box 59 1790 AB The Netherlands
2. School of Civil Engineering and Geosciences; Newcastle University; Newcastle-upon-Tyne NE1 7RU United Kingdom
3. Department of Earth Sciences, Faculty of Geosciences; Utrecht University; TA Utrecht P.O. Box 80.121, 3508 The Netherlands