1. Institute of Population Genetics, Universitätstrasse 1; Cluster of Excellence on Plant Sciences (CEPLAS), Heinrich Heine University; Universitätstrasse 40225 Düsseldorf Germany
2. Institute of Zoology, Department of Terrestrial Ecology, Zülpicher Str 47b; Cluster of Excellence on Plant Sciences (CEPLAS), University of Cologne; Zülpicher Strasse 50674 Köln Germany
3. Biodiversity and Climate Research Centre, Senckenberg Gesellschaft für Naturforschung, Senckenberganlage 25; 60325 Frankfurt am Main Germany