1. School of Environmental Sciences; University of East Anglia; Norwich Research Park Norwich NR4 7TJ UK
2. Department of Aquatic Geomicrobiology; Friedrich Schiller University Jena; Dornburger Str. 159 Jena 07743 Germany
3. School of Biological Sciences; University of East Anglia; Norwich Research Park Norwich NR4 7TJ UK
4. Plymouth Marine Laboratory; Prospect Place, The Hoe; Plymouth PL1 3DH UK
5. School of Life Sciences; University of Warwick; Coventry CV4 7AL UK