1. Department of Plant Protection, College of Plant Science and Technology; Huazhong Agricultural University; Wuhan 430070 China
2. The Key Lab of Crop Disease Monitoring & Safety Control in Hubei Province; Huazhong Agricultural University; Wuhan 430070 China
3. Department of Plant Pathology, College of Agriculture and Biotechnology; China Agricultural University; Beijing 100193 China
4. Department of Agricultural and Environmental Sciences; Clemson University; Clemson SC 29634 USA