1. Hawkesbury Institute for the Environment; University of Western Sydney; Building L9; Locked Bag 1797; Penrith South; NSW; 2751; Australia
2. School of Environmental Sciences; University of East Anglia; Norwich Research Park; Norwich; NR4 7TJ; UK
3. Landcare Research; PO Box 40; Lincoln; 7604; New Zealand
4. Institute of Biological and Environmental Sciences; University of Aberdeen; Zoology Building; Tillydrone Avenue; Aberdeen; AB24 2TZ; Scotland; UK