1. Forest Ecology and Forest Management Group Wageningen University and Research, P.O. Box 47 Wageningen 6700 AA The Netherlands
2. Department of Microbial Ecology Netherlands Institute of Ecology (NIOO‐KNAW) Wageningen 6708 PB The Netherlands
3. Ecology and Biodiversity Group, Department of Biology, Institute of Environmental Biology Utrecht University, Padualaan 8 Utrecht 3584 CH The Netherlands
4. Systems Ecology, Department of Ecological Science VU University (Vrije Universiteit) Amsterdam, De Boelelaan 1085 Amsterdam 1081 HV The Netherlands