1. Key Laboratory of Coastal Zone Environmental Processes and Ecological Remediation, Yantai Institute of Coastal Zone ResearchChinese Academy of Sciences Yantai 264003 China
2. University of Chinese Academy of Sciences Beijing 100049 China
3. Department of Gene EngineeringBinzhou Medical University Yantai 264003 China
4. Department of Marine SciencesUniversity of Connecticut Groton CT 06340 USA
5. State Key Laboratory of Estuarine and Coastal ResearchEast China Normal University Shanghai 200241 China