1. Key Laboratory of Beijing for Water Quality Science and Water Environmental Recovery Engineering; Engineering Research Center of Beijing; Beijing University of Technology; Beijing 100124 China
2. Advanced Water Management Centre (AWMC); The University of Queensland; St Lucia Brisbane QLD 4072 Australia
3. Microbiology, IWWR; Faculty of Science; Radboud University Nijmegen; Heyendaalseweg 135 6525 AJ Nijmegen The Netherlands
4. Department of Biochemistry and Microbiology; Laboratory of Microbiology; Gent University; Gent 9000 Belgium
5. Research Department of Microbiology; Beijing Genomics Institute (BGI)-Shenzhen; Shenzhen China