1. Helmholtz Centre for Environmental Research-UFZ, Department of Environmental Microbiology; Permoserstraße 15, 04318 Leipzig Germany
2. School of Engineering, Environmental Engineering, Newcastle University; Newcastle upon Tyne NE1 7RU UK
3. Department of Civil Engineering; University of Glasgow; Glasgow G12 8LT UK
4. Helmholtz Centre for Environmental Research-UFZ, Department of Ecological Modeling; Permoserstraße 15, 04318 Leipzig Germany