1. Institut National des Sciences Appliquées et de TechnologieUniversité Carthage Centre Urbain Nord, BP 676‐1080 Tunis Cedex Tunisia
2. Department of Molecular, Cellular, and Biomedical SciencesUniversity of New Hampshire Durham NH USA
3. Laboratoire Microorganismeset Biomolécules Actives (LR03ES03), Faculté des Sciences de TunisUniversité Tunis El Manar 2092 Tunis Tunisia
4. Hubbard Center for Genome StudiesUniversity of New Hampshire Durham NH USA
5. Department of Earth SciencesUniversity of New Hampshire Durham NH USA
6. Laboratory of Applied Microbiology, Department of Microbiology, Faculty of Natural and Life SciencesFerhat Abbas University 19000 Setif Algeria