1. Center for Marine Environmental Ecology School of Environmental Science and Engineering, Tianjin University Tianjin 300072 China
2. School of Marine and Atmospheric Sciences, Stony Brook University Stony Brook, New York 11794‐5000 USA
3. Marine Laboratory Duke University Beaufort NC 28516 USA
4. Biology Department Duke University Durham NC 27708 USA
5. State Key Laboratory of Marine Environmental Science College of Ocean and Earth Sciences, Xiamen University Xiamen Fujian 361005 China
6. Key Laboratory of Systems Bioengineering (Ministry of Education) Tianjin University Tianjin 300072 China