1. Laboratory of Microbiology Wageningen University & Research, Stippeneng 4 Wageningen 6708 WE The Netherlands
2. Department of Marine Microbiology and Biogeochemistry Royal Netherlands Institute for Sea Research, P.O. Box 59 Den Burg Texel 7197 AB The Netherlands
3. Centre of Biological Engineering University of Minho, Campus de Gualtar Braga 4710‐057 Portugal
4. Faculty of Geosciences Utrecht University, Princetonlaan 8a Utrecht 3584 CB The Netherlands