1. Biodiversity, University of Duisburg‐Essen Universitätsstr. 5, 45141 Essen Germany
2. Department of Ecology University of Kaiserslautern Erwin‐Schrödinger‐Str. 14, 67663 Kaiserslautern Germany
3. Lake and Glacier Research Institute of Ecology, University of Innsbruck Technikerstrasse 25, 6020 Innsbruck Austria