1. Geomicrobiology Group; College of Life Sciences; University of Dundee; Dundee DD1 5EH Scotland UK
2. The James Hutton Institute; Craigiebuckler Aberdeen AB15 8QH Scotland UK
3. Department of Soil and Environment; Swedish University of Agricultural Sciences; Uppsala Sweden
4. Laboratorio Biodiversità dei Funghi; Dipartimento di Biologia Ambientale; Sapienza Università di Roma; Rome 00185 Italy
5. Laboratory of Environmental Pollution and Bioremediation; Xinjiang Institute of Ecology and Geography; Chinese Academy of Sciences; Urumqi 830011 China