1. College of Marine Life Sciences, and Frontiers Science Center for Deep Ocean Multispheres and Earth System Ocean University of China, 5 Yushan Road Qingdao 266003 China
2. Laboratory for Marine Ecology and Environmental Science Qingdao National Laboratory for Marine Science and Technology Qingdao 266071 China
3. School of Biological Sciences University of East Anglia, Norwich Research Park Norwich NR47TJ UK
4. Institute of Evolution & Marine Biodiversity Ocean University of China Qingdao 266003 China