1. State Key Laboratory of Microbial Resources, Institute of Microbiology Chinese Academy of Sciences Beijing 100101 China
2. School of Environmental Sciences University of East Anglia, Norwich Research Park Norwich NR4 7TJ UK
3. School of Biological Sciences University of the Punjab Lahore 54590 Pakistan
4. State Key Laboratory of Soil and Sustainable Agriculture, Institute of Soil Science Chinese Academy of Sciences Nanjing Jiangsu Province 210008 China
5. College of Resources and Environment University of Chinese Academy of Sciences Beijing 10049 China
6. CAS Center for Excellence in Tibetan Plateau Earth Sciences Chinese Academy of Sciences Beijing 100101 China
7. Savaid Medical School University of the Chinese Academy of Sciences Beijing 10049 China