1. School of Biological Sciences; University of Essex; Essex CO4 3SQ UK
2. School of Geography, Earth and Environmental Sciences; University of Birmingham; Birmingham B15 2TT UK
3. Center for Environmental NanoScience and Risk, University of South Carolina; Columbia SC 29028 USA
4. Institute of Population Genetics, Cluster of Excellence on Plant Sciences (CEPLAS), Heinrich-Heine University; Düsseldorf 40225 Germany
5. Centre for Environment, Fisheries and Aquaculture Science; The Nothe, Barrack Road Weymouth Dorset DT4 8UB UK