1. State Key Laboratory of Lake Science and Environment; Nanjing Institute of Geography and Limnology, Chinese Academy of Sciences; Nanjing 210008 China
2. College of Resource and Environment, University of Chinese Academy of Sciences; Beijing 100049 China
3. Université Clermont Auvergne CNRS Laboratoire “Microorganismes: Génome et Environnement,”; CLERMONT-FERRAND F-63000 France