1. School of Biological Sciences; University of Reading; Reading RG6 6AJ UK
2. New Zealand Institute for Advanced Study; Massey University; Auckland New Zealand
3. Department of Environmental Microbiology; Eawag; Dübendorf 8600 Switzerland
4. Department of Environmental Systems Science; ETH Zürich; Zürich 8092 Switzerland
5. Max Planck Institute for Evolutionary Biology; Plön Germany