1. Department of Life Sciences; The Natural History Museum; London SW7 5BD UK
2. Cefas, Barrack Road, Weymouth; Dorset DT4 8UB UK
3. Department of Biological Sciences; University of Arkansas; Fayetteville AR 72701 USA
4. Department of Biological Sciences; Mississippi State University; MS State 39762 MS USA
5. Department of Ecology and Biodiversity; Utrecht University; 3584 Utrecht The Netherlands
6. Department of Terrestrial Ecology; Netherlands Institute of Ecology; 6708 PB Wageningen The Netherlands