1. School of Natural Resources and the Environment University of Arizona, Tucson, AZ, 85711, U.S.A.
2. U.S. Geological Survey Southwest Biological Science Center, Flagstaff, AZ, 86001, U.S.A.
3. South African National Biodiversity Institute Kirstenbosch Research Centre Claremont South Africa
4. Centre for Invasion Biology, Department of Botany and Zoology Stellenbosch University, Private Bag X1 Matieland 7602 South Africa