1. Ecology and Genetics Research Unit, Univ. of Oulu Oulu Finland
2. Dept of Biological and Environmental Sciences, Univ. of Gothenburg Gothenburg Sweden
3. CNRS, Univ. Lyon 1, Dept of Biometry and Evolutionary Biology, UMR 5558 Villeurbanne France
4. Rannevägen 12 Varberg Sweden