1. Graduate School of Integrated Arts and Sciences Kochi University Nankoku Japan
2. Faculty of Agriculture and Marine Science Kochi University Nankoku Japan
3. Kuroshio Zikkan Center Otsuki Japan
4. Fisheries Resources Institute, Japan Fisheries Research and Education Agency Sapporo Japan
5. Akkeshi Marine Station, Field Science Center for Northern Biosphere Hokkaido University Akkeshi Japan
6. The United Graduate School of Agricultural Science Ehime University Matsuyama Japan