1. Metapopulation Research Centre; Faculty of Biosciences; University of Helsinki; P.O. Box 65 (Viikinkaari 1) FI-00014 Helsinki Finland
2. Department of Life Sciences and Systems Biology; University of Turin; Via Accademia Albertina 13 10123 Torino Italy
3. Centre for Ecology, Evolution and Environmental Changes; Faculty of Sciences; University of Lisbon; Campo Grande; 1749-016 Lisbon Portugal
4. Faculty of Life Sciences; University of Madeira; Campus da Penteada; 9020-105 Funchal Portugal
5. Wildlife Research Unit; Laboratorio di Ecologia Applicata; Dipartimento di Agraria; Università degli Studi di Napoli Federico II; via Università 100, Portici Napoli 80055 Italy
6. School of Biological Sciences; University of Bristol; Bristol, 24 Tyndall Ave, Bristol BS8 1TH UK