1. Natural Resource Ecology Laboratory; Colorado State University; Fort Collins CO 80523 USA
2. Department of Biology; Colorado State University; Fort Collins CO 80523 USA
3. Hawkesbury Institute for the Environment and School of Science and Health; University of Western Sydney; Penrith NSW 2751 Australia
4. Faculty of Life Sciences, Michael Smith Building; The University of Manchester; Oxford Road Manchester M13 9PL UK
5. Department of Cell Biology, Microbiology and Molecular Biology; University of South Florida; Tampa FL 33620 USA