1. Centre for Biological Diversity; University of St Andrews; St Andrews Fife KY16 9TH United Kingdom
2. Institute of Environmental Sciences; Jagellonian University; Gronostajova 7 Kraków 30-387 Poland
3. Hawkesbury Institute for the Environment; University of Western Sydney; Penrith NSW 2751 Australia
4. Centre for Ecology and Conservation, College of Life and Environmental Sciences, University of Exeter; Cornwall Campus; Cornwall TR10 9EZ United Kingdom