1. Animal Breeding and Genomics Centre; Wageningen University; 6700 AH Wageningen the Netherlands
2. Research and Technology Centre; Hendrix Genetics; 5831 CK Boxmeer the Netherlands
3. Laboratory of Entomology; Wageningen University; 6700 EH Wageningen the Netherlands
4. Department of Terrestrial Ecology; Netherlands Institute of Ecology; 6700 AB Wageningen the Netherlands
5. Department of Biology; Indiana University; Bloomington Indiana 47405