1. Department of Ecology and Genetics; Evolutionary Biology Center and Science for Life Laboratory; Uppsala University; 75236 Uppsala Sweden
2. Institut des Sciences de l'Evolution (ISEM - UMR 5554 Université de Montpellier-CNRS-IRD-EPHE); Place Eugene Bataillon 34075 Montpellier France
3. Department of Ecology and Evolution; University of Toronto; 25 Willcocks St. Toronto ON M5S 3B2 Canada