1. Department of Science and Education Field Museum Chicago IL 60605 USA
2. Key Laboratory for Plant Diversity and Biogeography of East Asia Kunming Institute of Botany Chinese Academy of Sciences Kunming Yunnan 650201 China
3. Harvard University Herbaria Cambridge MA 02138 USA
4. Department of Botany California Academy of Sciences Golden Gate Park, San Francisco CA 94118 USA
5. Botanical Research Institute of Texas Fort Worth TX 76017 USA