1. Department of Ecology and Genetics, Plant Ecology and Evolution Uppsala University Norbyvägen 18D 75267 Uppsala Sweden
2. Department of Biology, The Bioinformatics Center University of Copenhagen Ole Maaløes Vej 5 2200 Copenhagen Denmark
3. Senckenberg Research Institute and Natural History Museum Senckenberganlage 25 60325 Frankfurt/Main Germany
4. Department of Botany University of Wyoming 1000 E. University Ave Laramie WY 82071 USA