1. School of Life Sciences; Sun Yat-Sen University; Guangzhou 510275 P. R. China
2. College of Animal Sciences and Technology; Sichuan Agricultural University; Ya'an 625014 Sichuan P. R. China
3. Key Laboratory of Tropical & Subtropical Fishery Resource Application & Cultivation; Ministry of Agriculture; Pearl River Fishery Research Institute; Chinese Academy of Fishery Sciences; Guangzhou 510380 P. R. China
4. Dazhong Breeding Co. Ltd; Jianyang 641400 P. R. China