1. School of Ecology and Environmental Sciences Yunnan University Kunming650091 YunnanChina
2. Yunnan Key Laboratory of Plant Reproductive Adaptation and Evolutionary Ecology Institute of Biodiversity Yunnan University Kunming 650504 Yunnan China
3. Environment Department Alcoa of Australia Ltd, Huntly Mine off Del Park Road PO Box 172 Pinjarra WA 6208 Australia