1. State Key Laboratory for Conservation and Utilization of Subtropical Agro‐Bioresources South China Agricultural University Guangzhou 510642 China
2. Guangdong Laboratory for Lingnan Modern Agriculture Guangzhou 510642 China
3. Guangdong Key Laboratory of New Technology in Rice Breeding Rice Research Institute Guangdong Academy of Agricultural Sciences Guangzhou 510640 China
4. Department of Cell and Developmental Biology John Innes Centre Norwich Research Park Norwich NR4 7UH UK