1. GEOMAR Helmholtz Center for Ocean Research; Marine Ecology/Food Webs; Duesternbrooker, Weg 20 D-24105 Kiel Germany
2. Institute of Marine Biology; National Taiwan Ocean University; 2 Pei-Ning Road Keelung 20224 Taiwan
3. Spanish Institute of Oceanography; IEO C.O. Santander; Promontorio de San Martín S/N 39004 Santander Spain
4. Université Lille, CNRS; Université du Littoral Côte d'Opale, UMR 8187; LOG, Laboratoire d'Océanologie et de Géosciences; F 62930 Wimereux France