1. Institut des Sciences de l’Évolution de Montpellier; CNRS (UMR 5554); Place Eugène Bataillon 34095 Montpellier cedex 05 France
2. Threatened Species Research Unit; South African National Biodiversity Institute; Private Bag X7 Kirstenbosch 7735 South Africa
3. Institute of Landscape and Plant Ecology; University of Hohenheim; 70593 Stuttgart Germany
4. Department of Biological Sciences; University of Cape Town; Private Bag; Rondebosch 7701 South Africa
5. Department of Molecular and Cell Biology; University of Cape Town; Rondebosch 7701 South Africa