1. Université de Bretagne Occidentale and Institut Universitaire Européen de la Mer; CNRS UMR 6538; Place Nicolas Copernic Plouzané Cedex 29280 France
2. National Institute of Polar Research; Tachikawa Tokyo 190-8518 Japan
3. Department of Polar Science; School of Multidisciplinary Science; Graduate University for Advanced Sciences; Tachikawa Tokyo 190-8518 Japan