1. Animal Genome Research Unit, Agrogenomics Research Center National Institute of Agrobiological Sciences Tsukuba Ibaraki Japan
2. Graduate School of Science and Engineering Yamagata University Yamagata Japan
3. Toxicology Division Institute of Environmental Toxicology Joso Ibaraki Japan
4. Technology and Development Team for Mouse Phenotype Analysis RIKEN BioResource Center Tsukuba Ibaraki Japan