1. National Key Laboratory of Wheat and Maize Crop Science/Collaborative Innovation Center of Henan Grain Crops/College of Agronomy Henan Agricultural University Zhengzhou 450002 China
2. Guangdong Provincial Key Laboratory for Plant Epigenetics, Longhua Institute of Innovative Biotechnology, College of Life Sciences and Oceanography Shenzhen University Shenzhen 518060 China
3. Hefei National Laboratory for Physical Sciences at the Microscale, Division of Life Sciences and Medicine, CAS Center for Excellence in Molecular Plant Sciences, School of Life Sciences University of Science and Technology of China Hefei 230027 China
4. Department of Biological Sciences and Biotechnology Research Center Michigan Technological University Houghton Michigan 49931 USA
5. College of Life Sciences Peking University Beijing 100871 China